Getting Started

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MCO invites applications from prospective students who have received a bachelor’s degree from an accredited college or university in the United States or its equivalent from an international institution.  First year graduate studies begin at the start of the fall semester, although it is possible to begin in July prior to the first year by undertaking a summer laboratory rotation.

Although both MCO and EPB are comprised of faculty from several departments, MCB is the administrative host of both programs.  When submitting an application, prospective students must indicate the program as well as department to which they are applying.

To apply to the Program in Molecules, Cells and Organisms, including the track in Engineering and Physical Biology, prospective students should complete the Graduate School of Arts and Sciences (GSAS) Online Application.  Paper applications will not be accepted.  In addition to being a training program comprised of faculty from five departments, The Program in Molecules, Cells and Organisms is a member of the Harvard Integrated Life Sciences (HILS) consortium, a federation of Harvard life sciences PhD programs, departments, and subject areas that facilitates cross-disciplinary academic and research collaboration.  See the HILS website (http://www.gsas.harvard.edu/hils/apply.php) for additional application details and information about the GSAS online application system.

Molecules, Cells and Organisms (MCO)

  • When filling out the GSAS Online Application, on the “Program Selection” page please indicate “Molecular and Cellular Biology” as your Program and “Molecules, Cells and Organisms” as your Subject.

Engineering and Physical Biology (EPB) Track in the MCO Program

  • When filling out the GSAS Online Application, on the “Program Selection” page please indicate “Molecular and Cellular Biology” as your Program and “Engineering and Physical Biology” as your Subject.